Usually means: Arrangement of multiple protein subunits.
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We found 8 dictionaries that define the word quaternary structure:

General (5 matching dictionaries)
  1. quaternary structure: American Heritage Dictionary of the English Language
  2. quaternary structure: Wiktionary
  3. quaternary structure: Dictionary.com
  4. Quaternary structure: Wikipedia, the Free Encyclopedia
  5. quaternary structure: TheFreeDictionary.com

Computing (1 matching dictionary)
  1. quaternary structure: Encyclopedia

Medicine (2 matching dictionaries)
  1. online medical dictionary (No longer online)
  2. quaternary structure: Medical dictionary

Definitions from Wiktionary (quaternary structure)

noun:  (biochemistry, molecular biology) The arrangement of multiple folded or coiled proteins to form a multi-subunit complex.

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