Usually means: Intermediate form between species evolution.
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We found 23 dictionaries that define the word missing link:

General (19 matching dictionaries)
  1. missing link: Longman Dictionary of Contemporary English
  2. missing link, the missing link: Merriam-Webster
  3. missing link: Oxford Learner's Dictionaries
  4. missing link: American Heritage Dictionary of the English Language
  5. missing link, the missing link: Collins English Dictionary
  6. missing link: Vocabulary.com
  7. missing link, the missing link: Cambridge Advanced Learner's Dictionary
  8. missing link: Wiktionary
  9. missing link: The Wordsmyth English Dictionary-Thesaurus
  10. missing link: Infoplease Dictionary
  11. missing link: Dictionary.com
  12. Missing Link (EP), Missing Link (TV series), Missing Link (band), Missing Link (film), Missing Link (puzzle), Missing Link, Missing link (disambiguation), Missing link (human evolution), Missing link, The Missing Link (Jeremy Enigk album), The Missing Link (Rage album), The Missing Link (wrestler), The Missing Link: Wikipedia, the Free Encyclopedia
  13. missing link: Rhymezone
  14. Missing Link: Dictionary of Phrase and Fable (1898)
  15. missing link: FreeDictionary.org
  16. missing link: Mnemonic Dictionary
  17. missing link: TheFreeDictionary.com

Computing (1 matching dictionary)
  1. The Missing Link, missing link: Encyclopedia

Miscellaneous (1 matching dictionary)
  1. missing link: The Skeptic's Dictionary

Slang (2 matching dictionaries)
  1. missing link: Green’s Dictionary of Slang
  2. The Missing Link: Urban Dictionary

(Note: See missing_links as well.)

Definitions from Wiktionary (missing link)

noun:  (colloquial) A hypothetical primate once thought necessary to explain a perceived evolutionary gap between apes and humans.
noun:  (figuratively) Any sought-after or valuable intermediary figure or position.

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