Might mean (unverified): Operating at full capacity efficiently.
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We found 2 dictionaries that define the word fire on all cylinders:

General (2 matching dictionaries)
  1. fire on all cylinders: Merriam-Webster
  2. fire on all cylinders: Wiktionary

Definitions from Wiktionary (fire on all cylinders)

verb:  (idiomatic) To operate as effectively as possible.

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