Usually means: Part of letter extending below baseline.
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We found 28 dictionaries that define the word descender:

General (18 matching dictionaries)
  1. descender: Merriam-Webster.com
  2. descender: Oxford Learner's Dictionaries
  3. descender: American Heritage Dictionary of the English Language
  4. descender: Collins English Dictionary
  5. descender: Vocabulary.com
  6. descender: Wordnik
  7. descender: Wiktionary
  8. descender: Webster's New World College Dictionary, 4th Ed.
  9. descender: Infoplease Dictionary
  10. Descender, descender: Dictionary.com
  11. Descender (disambiguation), Descender (typography), Descender: Wikipedia, the Free Encyclopedia
  12. Descender: Online Plain Text English Dictionary
  13. descender: Webster's Revised Unabridged, 1913 Edition
  14. descender: Rhymezone
  15. Descender, descender: AllWords.com Multi-Lingual Dictionary
  16. descender: Free Dictionary
  17. descender: Mnemonic Dictionary
  18. descender: Dictionary/thesaurus

Art (1 matching dictionary)
  1. ODLIS: Online Dictionary of Library and Information Science (No longer online)

Business (2 matching dictionaries)
  1. Bouvier's Law Dictionary 1856 Edition (No longer online)
  2. Descender: Legal dictionary

Computing (3 matching dictionaries)
  1. descender: Free On-line Dictionary of Computing
  2. descender: CCI Computer
  3. descender: Encyclopedia

Medicine (1 matching dictionary)
  1. online medical dictionary (No longer online)

Miscellaneous (1 matching dictionary)
  1. descender: A Word A Day

Sports (1 matching dictionary)
  1. CAVE AND KARST TERMINOLOGY (No longer online)

Tech (1 matching dictionary)
  1. Glossary of video terms (No longer online)

(Note: See descenders as well.)

Definitions from Wiktionary (descender)

noun:  A person or thing that descends.
noun:  (typography) The part of a letter or number that is drawn below the baseline (the bottom of most lowercase letters).
noun:  (typography, uncommon) A letter that has such a portion, such as g, j, or in some fonts Q.
noun:  (cycling) A cyclist who excels at fast descents.
noun:  (climbing) A belay device used in rock climbing, or as a fire escape.

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