Usually means: Beginning of any event or period.
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We found 11 dictionaries that define the word day one:

General (9 matching dictionaries)
  1. day one: Merriam-Webster.com
  2. day one: American Heritage Dictionary of the English Language
  3. day one: Collins English Dictionary
  4. day-one: Wordnik
  5. day one: Wiktionary
  6. day one: Infoplease Dictionary
  7. day one: Dictionary.com
  8. Day One (Birds of Tokyo album), Day One (From Ashes to New album), Day One (TV news series), Day One (TV program), Day One (TV series), Day One (Torchwood), Day One (app), Day One (band), Day One (building), Day one: Wikipedia, the Free Encyclopedia
  9. day one: Dictionary/thesaurus

Miscellaneous (1 matching dictionary)
  1. day one: Idioms

Slang (1 matching dictionary)
  1. Day One: Urban Dictionary

Definitions from Wiktionary (day one)

noun:  The very beginning.
noun:  (US, slang) A close, longtime friend, one who has been around since the beginning.
adverb:  At the very beginning; when something is entirely new.

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