Sorry, no online dictionaries contain the exact phrase branch on tree of life.
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Reverse dictionary results:
1. limb
2. stick
3. twig
4. graft
5. bush
6. rise
7. palm
8. knot
9. snag
10. deadwood
11. knotting
12. biology
13. silvics
14. arm
15. botany
16. son
17. perch
18. perched
19. perching
20. hamadryad
21. sap
22. leader
23. log
24. wand
25. wing
26. atm
27. ring
28. eden
29. girdle
30. bough
31. sticks
32. spur
33. plash
34. burl
35. grain
36. exobiology
37. caudex
38. olive branch
39. scrag
40. astrobiology

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You can look up the words in the phrase individually using these links:   branch ?   on ?   tree ?   of ?   life ?
(A question mark next to a word above means that we couldn't find it, but clicking the word might provide spelling suggestions.)

Not helpful? You might try using the wildcards * and ? to find the word you're looking for. For example, use
bran*to search for words beginning with bran, or
*lifeto search for words ending with life
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