Usually means: Species crucial for ecosystem balance.
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We found 9 dictionaries that define the word keystone species:

General (6 matching dictionaries)
  1. keystone species: Merriam-Webster
  2. keystone species: Oxford Learner's Dictionaries
  3. keystone species: Wiktionary
  4. keystone species: Dictionary.com
  5. Keystone species: Wikipedia, the Free Encyclopedia
  6. keystone species: TheFreeDictionary.com

Science (3 matching dictionaries)
  1. Environmental Terminology Discovery Service (No longer online)
  2. Glossary of Avian Terms For Use in Avian Conservation Biology (No longer online)
  3. Biodiversity (No longer online)

Definitions from Wiktionary (keystone species)

noun:  (ecology) A species that exerts a large, stabilizing influence throughout an ecological community, despite its relatively small numerical abundance.

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