Might mean (unverified): Joint connecting vertebrae in reptiles.
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We found 2 dictionaries that define the word zygosphere:

General (2 matching dictionaries)
  1. zygosphere: Merriam-Webster
  2. zygosphere: Wiktionary

Definitions from Wiktionary (zygosphere)

noun:  (botany) A plant's gamete which can form a zygospore by uniting with a similar one.

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