Usually means: Relating to bats, typically nocturnal.
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We found 3 dictionaries that define the word vespertilian:

General (3 matching dictionaries)
  1. vespertilian: Merriam-Webster
  2. vespertilian: Collins English Dictionary
  3. vespertilian: Wiktionary

Definitions from Wiktionary (vespertilian)

adjective:  of, pertaining to, or characteristic of bats.

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