Usually means: Air-filled cavity in plants.
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We found 3 dictionaries that define the word pneumatode:

General (2 matching dictionaries)
  1. pneumatode: Merriam-Webster
  2. pneumatode: Wiktionary

Science (1 matching dictionary)
  1. Bryological (No longer online)

Definitions from Wiktionary (pneumatode)

noun:  (botany, histology) A group of cells with spiral secondary wall thickenings, present in a velamen; acts as an airway for gas exchange for respiration or photosynthesis
noun:  (botany, histology) A group of cells presumed to be specialised as an airway for ventilation of particular tissues.

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