Usually means: Leaf surface ecosystem for microorganisms.
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We found 5 dictionaries that define the word phyllosphere:

General (4 matching dictionaries)
  1. phyllosphere: Collins English Dictionary
  2. phyllosphere: Wiktionary
  3. Phyllosphere: Wikipedia, the Free Encyclopedia
  4. phyllosphere: TheFreeDictionary.com

Science (1 matching dictionary)
  1. Agricultural Thesaurus and Glossary (No longer online)

Definitions from Wiktionary (phyllosphere)

noun:  (biology) The above-ground surface of plants, viewed as a habitat for microorganisms

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