Usually means: Group of hoofed African mammals.
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We found 3 dictionaries that define the word paenungulate:

General (3 matching dictionaries)
  1. paenungulate: Merriam-Webster
  2. paenungulate: Wiktionary
  3. Paenungulate: Wikipedia, the Free Encyclopedia

Definitions from Wiktionary (paenungulate)

noun:  (zoology) Any animal of the clade Paenungulata.

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