Usually means: Respiratory system with one spiracle.
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We found 4 dictionaries that define the word metapneustic:

General (4 matching dictionaries)
  1. metapneustic: Merriam-Webster
  2. metapneustic: Wordnik
  3. metapneustic: Wiktionary
  4. metapneustic: Grandiloquent Dictionary

Definitions from Wiktionary (metapneustic)

adjective:  (zootomy) Having the respiratory apparatus in the anus.

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