Usually means: Largest chamber in some foraminifera.
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We found 4 dictionaries that define the word megalosphere:

General (3 matching dictionaries)
  1. megalosphere: Merriam-Webster
  2. megalosphere: Wordnik
  3. megalosphere: Wiktionary

Computing (1 matching dictionary)
  1. megalosphere: Encyclopedia

Definitions from Wiktionary (megalosphere)

noun:  A foraminifer that has a large initial chamber

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