Usually means: Cluster of microspores in ferns.
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We found 4 dictionaries that define the word massula:

General (3 matching dictionaries)
  1. massula: Merriam-Webster
  2. massula: Wordnik
  3. massula: Wiktionary

Science (1 matching dictionary)
  1. Botanical Terms (No longer online)

Definitions from Wiktionary (massula)

noun:  (palynology) A union of pollen microspores.

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