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We found 2 dictionaries that define the word in copula:

General (1 matching dictionary)
  1. in copula: Wiktionary

Science (1 matching dictionary)
  1. DRAGONFLY TERMS (No longer online)

Definitions from Wiktionary (in copula)

adjective:  (zoology) During copulation; in the copulative state.
adverb:  (zoology) During copulation; in the copulative state.

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