Might mean (unverified): Joint evolutionary history of species.
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We found 2 dictionaries that define the word cospeciation:

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  1. cospeciation: Wiktionary
  2. Cospeciation: Wikipedia, the Free Encyclopedia

Definitions from Wiktionary (cospeciation)

noun:  An event that occurs when a symbiotic relationship between two organisms leads to a change creating a new species in the process.

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