Sorry, no online dictionaries contain the exact phrase be in on the joke.
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Reverse dictionary results:
1. in on gag
2. in on prank
3. in on fun
4. in on secret
5. in on laugh
6. comedy
7. farce
8. fun
9. funny
10. gag
11. history
12. humor
13. humour
14. jape
15. jest
16. joked
17. jokes
18. joking
19. kid
20. kidding
21. laugh
22. laughing
23. laughs
24. laughter
25. pun
26. ridicule
27. serious
28. smile
29. acting
30. amusement
31. caprice
32. kicks
33. kif
34. know
35. practical
36. rib
37. sally
38. seriously
39. shenanigan
40. something

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You can look up the words in the phrase individually using these links:   be ?   in ?   on ?   the ?   joke ?
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Not helpful? You might try using the wildcards * and ? to find the word you're looking for. For example, use
be i*to search for words beginning with be i, or
*joketo search for words ending with joke
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